The exact formula is currently unknown, but it depends on the flatness of and amount of water on tiles in the central 25 with a Manhattan distance of 3 or less from the start square (including the start square itself). The population requirement to reach a milestone varies depending on the map selected. You can have a total of nine areas unlocked by the end of the game, but this could be increased to a total of 25 areas or even 81 areas with Steam Workshop modifications. Some of the areas can have water areas, highways, and/or natural resources.
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